Penny Gullan

B.Sc. (Honours) Zoology, Monash University, Australia
Ph.D. Entomology, Monash University, Australia 


Systematics (taxonomy and phylogeny) and biology of scale insects (Hemiptera: Coccoidea), especially soft scales, eriococcids, mealybugs and archaeococcoids; coccoid-plant interactions, especially involving gall-inducing taxa.

 

Publications

Books and articles in refereed journals

  1. Kondo, T., Ramos-Portilla, A.A., Peronti, A.L.B.G. and Gullan, P.J. (2016) Known distribution and pest status of fluted scale insects (Hemiptera Monophlebidae Iceryini) in South America. Redia 94: 187–195. pdf
  2. Gullan, P.J. and Williams, D.J. (2016) A new pupillarial scale insect (Hemiptera: Coccoidea: Eriococcidae) from Angophora in coastal New South Wales, Australia. Zootaxa 4117(1): 85–100. [Free PDF available from journal at: http://www.mapress.com/j/zt/article/view/zootaxa.4117.1.4]
  3. Kondo, T., Gullan, P.J. and Cook. L.G. (2016) A review of the genus Capulinia Signoret (Hemiptera: Coccoidea: Eriococcidae) with description of two new species. Zootaxa 4111(4): 471–491. [Free PDF available from journal at: http://www.mapress.com/j/zt/article/view/zootaxa.4111.4.7]
  4. Kondo, T., Gullan, P.J., Peronti, A.L.B.G., Ramos-Portilla, A.A., Caballero, A. and Villarreal, N. (2016) First records of the iceryine scale insects Crypticerya brasiliensis (Hempel) and Crypticerya genistae (Hempel) (Hemiptera: Monophlebidae) for Colombia. Insecta Mundi 0480: 1–9. pdf
  5. Neumann, G., O'Dowd, D.J., Gullan, P.J. and Green, P.T. (2016) Diversity, endemism and origins of scale insects on a tropical oceanic island: implications for management of an invasive ant. Journal of Asia-Pacific Entomology 19: 159–166. pdf
  6. Semple, T.L., Gullan, P.J., Hodgson, C.J., Hardy, N.B. and Cook, L.G. (2015) Systematic review of the Australian 'bush coconut' genus Cystococcus (Hemiptera: Eriococcidae) uncovers a new species from Queensland. Invertebrate Systematics 29: 287–312. [Free PDF available from journal at: http://www.publish.csiro.au/paper/IS14061.htm]
  7. Kondo, T., Gullan, P.J., Watason, G.W., Bustillo Pardey, A.E. and Montes, L.G. (2015) New distribution and host records for white coconut scale, Parlagena bennetti Williams (Hemiptera: Diaspididae). Insecta Mundi 0422: 1–6. pdf
  8. Lin, Y.-P, Cook, D.H., Gullan, P.J. and Cook, L.G. (2015) Does host-plant diversity explain species richness in insects? A test using Coccidae (Hemiptera). Ecological Entomology 40: 299–306. pdf
  9. Foldi, I. and Gullan, P.J. (2014) Review of the Neotropical scale insects formerly assigned to Coelostomidiidae and here transferred to a new tribe within the Monophlebidae (Hemiptera: Sternorrhyncha: Coccoidea). Zootaxa 3903 (1): 1–62. [Free PDF available from Zootaxa: <http://www.mapress.com/zootaxa/2014/f/zt03903p062.pdf>]
  10. Gullan, P.J. and Cranston, P.S. (2014) The Insects: An Outline of Entomology. 5th edition. Wiley-Blackwell, 595 pp.
  11. Neumann, G., O'Dowd, D.J., Gullan, P.J. and Green, P.T. (2014) First record of Pulvinaria urbicola Cockerell (Hemiptera: Coccidae), a potentially damaging scale insect, on Christmas Island, Indian Ocean. Journal of Asia-Pacific Entomology 17: 27–30. pdf
  12. Mech, A.M., Asaro, C., Cram, M.M., Coyle, D.R., Gullan, P.J., Cook, L.G. and Gandhi, K.J.K. (2013) Matsucoccus macrocicatrices (Hemiptera: Matsucoccidae): first report, distribution, and association with symptomatic eastern white pine in the southeastern United States. Journal of Economic Entomology 106: 2391–2398. pdf
  13. Gullan, P.J., Moir, M.L. and Leng, M.C. (2013) A new species of mealybug (Hemiptera: Pseudococcidae) from critically endangered Banksia montana in Western Australia. Records of the Western Australian Museum 28: 13–20. pdf
  14. Wu, S.A., Nan, N., Gullan, P. and Deng, J. (2013) The taxonomy of the Japanese oak red scale insect, Kuwania quercus (Kuwana) (Hemiptera: Coccoidea: Kuwaniidae), with a generic diagnosis, a key to species and description of a new species from California. Zootaxa 3630(2): 291–307. pdf
  15. Lin, Y.-P., Kondo, T., Gullan, P. and Cook, L.G. (2013) Delimiting genera of scale insects: molecular and morphological evidence for synonymising Taiwansaissetia Tao, Wong and Chang with Coccus Linnaeus (Hemiptera: Coccoidea: Coccidae). Systematic Entomology 38: 249–264. DOI: 10.1111/j.1365-3113.2012.00664.x
  16. Kondo, T., Gullan, P. and Ramos Portilla, A.A. (2012) Report of new invasive scale insects (Hemiptera; Coccoidea), Crypticerya multicicatrices Kondo and Unruh (Monophlebidae) and Maconellicoccus hirsutus (Green) (Pseudococcidae), on the islands of San Andres and Providencia, Colombia, with an updated taxonomic key to iceryine scale insects of South America. Insecta Mundi 0265: 1–17. pdf
  17. Kaydan, M.B. and Gullan, P.J. (2012) A taxonomic revision of the mealybug genus Ferrisia Fullaway (Hemiptera: Pseudococcidae), with descriptions of eight new species and a new genus. Zootaxa 3543: 1–65. [Free PDF available from Zootaxa: <http://www.mapress.com/zootaxa/2012/f/zt03543p065.pdf>]
  18. Hodgson, C.J., Millar, I.M. and Gullan, P.J. (2011) Cissococcus Cockerell (Hemiptera: Coccoidea: Coccidae), a unique gall-inducing soft scale genus on Vitaceae from South Africa, with description of a new species. Zootaxa. 2996: 1–32. pdf
  19. Hardy, N.B., Beardsley, J.W. and Gullan, P.J. (2011) Uncovering diversity of Australian Eucalyptus-constrained felt scales (Hemiptera: Coccoidea: Eriococcidae). Systematic Entomology 36: 497-528. pdf
  20. Kondo, T. and Gullan, P.J.  (2011) Taxonomic review of the genus Tachardiella Cockerell, with a key to species of lac insects (Hemiptera: Coccoidea: Kerriidae) recorded from the New World. Neotropical Entomology. pdf
  21. Williams, D.J., Gullan, P.J., Miller, D.R., Matile-Ferrero, D. and Han, S.I. (2011) A study of the scale insect genera Puto Signoret (Hemiptera: Sternorrhyncha: Coccoidea: Putoidae) and Ceroputo Sulc (Pseudococcidae), with a comparison to Phenacoccus (Pseudococcidae). Zootaxa 2802: 1-22. pdf
  22. Kondo, T., Gullan, P.J. and Pemberton, R.W. (2011) A new species of Paratachardina Balachowsky (Hemiptera: Coccoidea: Kerriidae) related to the pestiferous lobate lac insect P. pseudolobata Kondo & Gullan. Journal of Asia-Pacific Entomology 14: 141-146. pdf
  23. Gruwell, M.E., Hardy, N.B., Gullan, P.J. and Dittmar, K. (2010) Evolutionary relationships among primary endosymbionts of the mealybug subfamily Phenacoccinae (Hemiptera: Coccoidea: Pseudococcidae). Applied and Environmental Microbiology 76(22): 7521-7525. Doi:10.1128/AEM.01354-10
  24. Lin, Y.-P., Gullan, P.J. and Cook, L.G. (2010) Species richness and host-plant diversity are positvely correlated in Coccidae. Entomologia Hellenica 19: 90-98. pdf
  25. Hardy, N.B. and Gullan, P.J. (2010) Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea: Eriococcidae) and descriptions of a new genus and nine new species. ZooKeys 58: 1-74.pfd
  26. Kondo, T. and Gullan, P.J. (2010) The Coccidae (Hemiptera: Coccoidea) of Chile, with descriptions of three new species and transfer of Lecanium resinatum Kieffer & Herbst to the Kerriidae. Zootaxa 2560: 1-15. [Free PDF available from Zootaxa]
  27. Williams, D.J. and Gullan, P.J. (2010) Family-group names proposed in the family Pseudococcidae (Hemiptera: Sternorrhyncha: Coccoidea). Zootaxa 2400: 66-68.
  28. Gullan, P.J. Kaydan, B. M. & Hardy, N.B.(2010) Molecular phylogeny and species recognition in the mealybug genus Ferrisia Fullaway (Hemiptera: Pseudococcidae). Systematic Entomology 35: 329–339. pfd (corrected proof)
  29. Gullan, P.J. and Cranston, P.S. (2010) The Insects: An Outline of Entomology. 4th edition. Wiley-Blackwell, 565 pp.
  30. Gullan, P.J. and Martin, J.H. (2009) Sternorrhyncha (jumping plant-lice, whiteflies, aphids, and scale insects). pp. 957–967. In: Resh, V.H. & Cardé, R.T. (eds.) Encyclopedia of Insects. 2nd edn. Elsevier, San Diego.pdf
    [The attached PDF is a preprint of the chapter (with some uncorrected errors). Please see the publisher’s web site for information on the Encyclopedia of Insects, at: http://www.elsevier.com/wps/find/bookdescription.cws_home/718546/descrip...
  31. Cranston, P.S. and Gullan, P.J. (2009) Phylogeny of insects. pp. 780–793. In: Resh, V.H. & Cardé, R.T. (eds.) Encyclopedia of Insects. 2nd edn. Elsevier, San Diego. pdf The attached PDF is a preprint of the chapter (with some uncorrected errors). Please see the publisher’s web site for information on the Encyclopedia of Insects, at: http://www.elsevier.com/wps/find/bookdescription.cws_home/718546/descrip...
  32. Gullan, P.J. and Williams, D.J. (2009) Chris J. Hodgson – much more than an expert coccidologist. pp. 315–317. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal. pdf
  33. Gullan, P.J. (2009) Jan Koteja – an extraordinary evolutionary and revolutionary coccidologist. pp. 305–307. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal. pdf
  34. Gullan, P.J. and Kondo, T. (2009) The morphology of lac insects (Hemiptera: Coccoidea: Kerriidae). pp. 63–70. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal. pdf
  35. Hardy, N.B. and Gullan, P.J. (2009) Opisthoscelis nigra Froggatt (Eriococcidae) is an armoured scale Maskellia nigra (Froggatt) (Diaspididae). pp. 59–62. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal. pdf
  36. Kondo, T., Gullan, P.J. and Williams, D.J. (2008) The study of scale insects (Hemiptera: Sternorrhyncha: Coccoidea). Revista Corpoica - Ciencia y Tecnologia Agropecuaria 9(2): 55–61. pdf
  37. Schroer, S., Pemberton, R.W., Cook, L.G., Kondo, T. and Gullan, P.J. (2008) The genetic diversity, relationships, and potential for biological control of the lobate lac scale, Paratachardina pseudolobata Kondo & Gullan (Hemiptera: Coccoidea: Kerriidae). Biological Control 46: 256–266.pdf
  38. Oliveira, C.M., Frizzas, M.R., Fialho, J.F. and Gullan, P.J. (2008) Biology of Protortonia navesi (Hemiptera: Monophlebidae), a new cassava pest in Brazil, with notes on its behavior in the field. Annals of the Entomological Society of America 101: 779–785. pdf
  39. Hardy, N.B., Gullan, P.J., Henderson, R.C. and Cook, L.G. (2008) Relationships among scale insects (Hemiptera: Coccoidea: Eriococcidae) of southern beech, Nothofagus (Nothofagaceae), with the first descriptions of Australian species of the Nothofagus-feeding genus Madarococcus Hoy. Invertebrate Systematics 22: 365–405. pdf[Please note that CSIRO Publishing only allows a pre-publication version of this article to be posted here; please go to the home page of the journal Invertebrate Systematics for the final version: http://www.publish.csiro.au/nid/120.htm or contact the authors directly]
  40. Unruh, C.M. and Gullan, P.J. (2008) Identification guide to species in the scale insect tribe Iceryini (Coccoidea: Monophlebidae). Zootaxa 1803: 1–106. pdf
  41. Williams, D.J. and Gullan, P.J. (2008) A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini), Journal of Natural History 42: 77–128.M pdf
  42. Kondo, T. and Gullan, P.J. (2008) Synonymy of Plotococcus Miller & Denno with Leptococcus Reyne, and description of a new species from Colombia (Hemiptera: Pseudococcidae). Neotropical Entomology 37: 51–57. pdf
  43. Hardy, N.B. and Gullan, P.J. (2008) Case number 3424. Pseudococcini Cockerell, 1905 (insecta, Hemiptera): proposed precedence over Sphaerococcini Cockerell, 1899. Bulletin of Zoological Nomenclature 64(1): 1–3.pdf and the 2009 judgement (opinion)! pdf
  44. Cook, L.G. and Gullan, P.J. (2008) Insect, not plant, determines gall morphology in the Apiomorpha pharetrata species-group (Hemiptera: Coccoidea). Australian Journal of Entomology 47: 51–57. pfd
  45. Hardy, N.B., Gullan, P.J. and Hodgson, C.J. (2008) A subfamily-level classification of mealybugs (Hemiptera: Pseudococcidae) based on integrated molecular and morphological data. Systematic Entomology 33: 51–71. pfd
  46. Unruh, C.M. and Gullan P. J. (2008) Molecular data reveal convergent reproductive strategies in iceryine scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing re-interpretation of morphology and a revised generic classification. Systematic Entomology 33: 8–50. pdf
  47. Gullan, P.J. and Cook, L.G. (2007) Phylogeny and higher classification of the scale insects (Hemiptera: Sternorrhyncha: Coccoidea). In Zhang, Z.-Q. & Shear, W.A. (Eds) Linnaeus Tercentenary: Progress in Invertebrate Taxonomy. Zootaxa 1668: 413–425. pdf
  48. Culik, M.P., Martins, D.S., Ventura, J.A., Peronti, A.L.B.G., Gullan, P.J. and Kondo, T. (2007) Coccidae, Pseudococcidae, Ortheziidae, and Monophlebidae (Hemiptera: Coccoidea) of Espérito Santo, Brazil. Biota Neotropica 7(3): 61–65.pdf
  49. 84. Kondo, T. and Gullan, P.J. (2007) Taxonomic review of the lac insect genus Paratachardina Balachowsky (Hemiptera: Coccoidea: Kerriidae), with a revised key to genera of Kerriidae and description of two new species. Zootaxa 1617: 1–41. pdf
  50. Dorchin, N. and Gullan, P.J. (2007) A new genus and species of a lasiopterine gall midge (Diptera: Cecidomyiidae) from bud galls on renosterbos, Elytropappus rhinocerotis (Asteraceae), in South Africa. African Entomology 15: 233–240.
  51. Hardy, N.B. and Gullan, P.J. A new species of Phacelococcus Miller (Hemiptera: Coccoidea: Eriococcidae) on Bursaria (Pittosporaceae). The Australian Entomologist 34: 85–91.
  52. Hardy, N.B. and Gullan, P.J. (2007) A new genus and four new species of felt scales on Eucalyptus (Hemiptera: Coccoidea: Eriococcidae) in southeastern Australia. Australian Journal of Entomology 46: 106–120. pdf
  53. Kondo, T., Hardy, N., Cook, L. and Gullan, P. (2006) Descriptions of two new genera and species of Eriococcidae (Hemiptera: Coccoidea) from southern South America. Zootaxa 1349: 19–36. pdf
  54. Unruh, C.M. and Gullan, P.J. (2007) Hypogeal margarodids of the genus Heteromargarodes Jakubski (Coccoidea: Margarodidae) from the western United States. Proceedings of the Entomological Society of Washington 109: 166–181. pdf
  55. Booth, J.M. and Gullan, P.J. (2006) Synonymy of three pestiferous Matsucoccus scale insects (Coccoidea: Margarodidae) based on morphological and molecular evidence. Proceedings of the Entomological Society of Washington 108: 749–760. pdf
  56. Williams, D.J., Gullan, P.J., Englberger, K. and Moore, A. (2006) Report on the scale insect Icerya imperatae Rao (Hemiptera: Coccoidea: Margarodidae) seriously infesting grasses in the Republic of Palau. Micronesica 38(2): 267–272.pdf
  57. Culik, M.P., Martins, D. dos S. and Gullan, P.J. (2006) First records of two mealybug species in Brazil and new potential pests of papaya and coffee. Journal of Insect Science 6 (23): 1–6. pdf
  58. Gullan, P.J., Giliomee, J.H., Hodgson, C.J. and Cook, L.G. (2006) The systematics and biology of the South African gall-inducing scale insect Calycicoccus merwei Brain (Hemiptera: Coccoidea: Eriococcidae). African Entomology 14: 13–33.pdf
  59. Kondo, T., Gullan, P.J., Ventura, J.A. and Culik, M. (2005)Taxonomy and biology of the mealybug genus Plotococcus Miller & Denno (Hemiptera: Pseudococcidae) in Brazil, with descriptions of two new species. Studies on Neotropical Fauna and Environment  40: 213–227. pdf
  60. Kondo, T., Williams, M.L. and Gullan, P.J. (2005) Redescription of Octolecanium perconvexum (Cockerell), new genus and new combination, with description of new species from Guatemala (Hemiptera: Coccoidea: Coccidae). TIP Revista Especializada en Ciencias Químico-Biológicas 8(1): 5–11.pdf
  61. Kondo, T. and Gullan, P.J. (2005) A new species of Austrotachardiella Chamberlin on guava in Colombia (Hemiptera: Coccoidea: Kerriidae), with a revised key to the species of the genus. Neotropical Entomology 34(3): 401–407. pdf
  62. Culik, M.P. and Gullan, P.J. (2005) A new pest of tomato and other records of mealybugs (Hemiptera: Pseudococcidae) from Espírito Santo, Brazil. Zootaxa 964: 1–8. pdf
  63. Gullan, P.J., Miller, D.R. and Cook. L.G. (2005) Gall-inducing scale insects (Hemiptera: Sternorrhyncha: Coccoidea). pp. 159–229. In: A. Raman, C.W. Schaefer and T.M. Withers (eds), Biology, Ecology, and Evolution of Gall-Inducing Arthropods. Science Publishers, Inc., Enfield, USA, Plymouth, UK. pdf-part1(7mb) pdf-part2 (4mb) refs (396k)
  64. Kondo, T., Williams, M. L. and Gullan, P.J. (2005) Taxonomic review of the genus Xenolecanium Takahashi and description of the new genus Takahashilecanium Kondo (Hemiptera: Coccidae; Coccinae, Paralecaniini). Entomological Science 8: 109–120. pdf
  65. Downie, D.A. and Gullan, P.J. (2005) Phylogenetic congruence of mealybugs and their primary endosymbionts. Journal of Evolutionary Biology 18: 31–324. pdf
  66. Gullan, P.J. and Cranston, P.S. (2005) The Insects: An Outline of Entomology. 3rd edition. Blackwell Publishing Ltd, 505 pp.
  67. Kondo, T., Gullan, P.J. and Miller, D.R. (2004) A new hypogeal species of Oregmopyga Hoy (Hemiptera: Coccoidea: Eriococcidae) from southern California, U.S.A., and a key to species. Zootaxa 784: 1–12. pdf
  68. Cook, L.G. and Gullan, P.J. (2004) Gall induction has evolved multiple times among the eriococcid scale insects (Sternorrhyncha: Coccoidea: Eriococcidae). Biological Journal of the Linnean Society 83: 441–452. pdf
  69. Austin, A.D., Yeates, D.K., Cassis, G., Fletcher, M.J., La Salle, J., Lawrence, J.F., McQuillan, P.B., Mound, L.A., Bickel, D.J., Gullan, P.J., Hales, D.F. and Taylor, G.S. (2004) Insects 'down under' - diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43: 216–234.pdf
  70. Kondo, T. and Gullan, P.J. (2004) A new species of ant-tended soft scale of the genus Cryptostigma Ferris (Hemiptera: Coccidae) associated with bamboo in Peru. Neotropical Entomology 33: 717–723. pdf
  71. Laflin, H.M., Gullan, P.J. and Parrella, M.P. (2004) Mealybug species (Pseudococcidae) found on ornamental crops in California nursery production. Proceedings of the Entomological Society of Washington 106: 475–477. pdf
  72. Downie, D.A. and Gullan, P.J. (2004) Phylogenetic analysis of mealybugs (Hemiptera: Pseudococcidae) based on DNA sequences from three nuclear genes, and a review of the higher classification. Systematic Entomology 29: 238–259. pdf
  73. Gullan, P.J., Downie, D.A. and S.A. Steffan (2003) A new pest species of the mealybug genus Ferrisia Fullaway (Hemiptera: Pseudococcidae) from the United States. Annals of the Entomological Society of America 96: 723–737. pdf
  74. Thao, M.L., Baumann, L., Hess, J.M., Falk, B.W., Ng, C.K., Gullan, P.J. and Baumann, P. (2003) Phylogenetic evidence for two new insect-associated chlamydia of the family Simkaniaceae. Current Microbiology 47: 46–50. pdf
  75. Cranston, P.S. and Gullan, P.J. (2003) Phylogeny of insects. pp. 882-898. In V.H. Resh and R. T. Cardé (eds), Encyclopedia of Insects. Academic Press. Amsterdam. pdf
  76. 57. Gullan, P.J. and Martin, J.H. (2003) Sternorrhyncha (jumping plant-lice, whiteflies, aphids and scale insects). pp.1079–1089. In V. Resh and R. Cardé (eds), Encyclopedia of Insects. Academic Press Amsterdam. pdf
  77. Cook, L.G., Gullan, P.J. and Trueman, H.E. (2002) A preliminary phylogeny of the scale insects (Hemiptera: Sternorrhyncha: Coccoidea) based on nuclear small-subunit ribosomal DNA. Molecular Phylogenetics and Evolution 25: 43–52. pdf
  78. Thao, M.L., Gullan, P.J. and Baumann, P. (2002) Secondary (-Proteobacteria) endosymbionts infect the primary (β-Proteobacteria) endosymbionts of mealybugs multiple times and coevolve with their hosts. Applied and Environmental Microbiology 68(7): 3190–3197. pdf
  79. Cook, L.G. and Gullan, P.J. (2002) Longevity and reproduction in ApiomorphaRübsaamen (Hemiptera: Sternorrhyncha: Coccoidea). Bollettinodi Zoologia agraria e di Bachicoltura,series II, 33(3): 259–265. pdf
  80. Gullan, P.J. and Cook, L.G. (2002) Phenacoleachia Cockerell (Hemiptera:Coccoidea: Phenacoleachiidae) re-visited. Bollettinodi Zoologia agraria e di Bachicoltura,series II, 33(3): 163–173. pdf
  81. Gullan, P.J. and Sjaarda, A.W. (2001) Trans-Tasman Platycoelostoma Morrison (Hemiptera: Coccoidea: Margarodidae) on endemic Cupressaceae, and the phylogenetic history of margarodids. Systematic Entomology 26: 1–22.pdf
  82. Gullan, P.J. (2001) Why the taxon Homoptera does not exist. Entomologica 33 (1999): 101–104. pdf
  83. Lit, I.L., Jr and Gullan, P.J. (2001) Comparative morphology of the analtubercle and associated structures of some lac insects (Kerriidae, Coccoidea,Hemiptera). Entomologica 33 (1999): 119–126.
  84. Gullan, P.J. and Cook, L.G. (2001) Are cochineal insects eriococcids? Entomologica 33 (1999): 91–9. pdf
  85. Cook, L.G. and Gullan, P.J. (2001) Are the enlarged ducts of Eriococcus (Coccoidea: Eriococcidae) plesiomorphic? Entomologica 33 (1999): 59–66.
  86. Gullan, P.J. (2000) Identification of the immature instars of mealybugs (Hemiptera: Pseudococcidae) found on citrus in Australia. Australian Journal of Entomology 39: 160–166. pdf
  87. Cook, L.G., Gullan, P.J.,Stewart, A.C. (2000) First-instar morphology and sexual dimorphism in the gall-inducing scale insect Apiomorpha (Hemiptera: Coccoidea: Eriococcidae). Journal of Natural History. 34: 879–894. pdf
  88. Gullan, P.J. and Cranston, P.S. (2000) The Insects: An Outline of Entomology. 2nd edition. Blackwell Science, Oxford. 470 pp. + appendix.
  89. Qin, T.K. and Gullan, P.J. (1999) A new synonym of Ceroplastes destructor Newstead (Hemiptera: Coccoidea: Coccidae: Ceroplastinae). African Entomology 7: 305–306. pdf
  90. Gullan, P.J. (1999) A new genus of subcortical coccoids (Hemiptera: Eriococcidae). Memoirs of the Museum of Victoria. 57: 241–250. pdf
  91. Heckroth, H.-P., Fiala, B., Gullan, P.J. et al. (1998) The soft scale (Coccoidae) associates of Malaysian ant-plants. Journal of Tropical Ecology 14: 427–444. pdf
  92. Gullan, P.J. and Brookes, H.M. (1998) Giant females and bird-of-paradise flies: notes on the biology of Callipappus Guérin-Méneville (Hemiptera: Margarodidae). Australian Journal of Entomology. 37: 2–7.
  93. Gullan, P.J. and Hodgson, C.J. (1998) A new genus of Australian soft scale insect (Hemiptera: Coccidae) with species on Capparis (Capparaceae) and Doryphora (Monimiaceae) from New South Wales. Proceedings of the Linnean Society of New South Wales 119: 197–217.
  94. Miller, D.R., Gullan, P.J. and Williams, D.J. (1998) Family placement of species previously included in the scale insect genus Sphaerococcus Maskell (Hemiptera; Cocoidea). Proceedings of the Entomological Society of Washington  100: 286–303.
  95. Qin, T.K. and Gullan, P.J. (1998) Systematics as a tool for pest management: case studies using scale insects and mites. Proceedings of the Sixth Australasian Applied Entomological Research Conference, Brisbane, 29 Sept - 2 Oct. 1998, vol 1: 479–488.
  96. Qin, T.K., Gullan, P.J. and G.A.C. Beattie (1998) Biogeography of the wax scales (Insecta: Hemiptera: Coccidae: Ceroplastinae). Journal of Biogeography. 25: 37–45. pdf
  97. Gullan, P.J. (1997) Relationships with ants. Ch. 1.4.8. pp. 351–373. In: Ben-Dov, Y. and Hodgson, C.J. (Eds). Soft Scale Insects - Their Biology, Natural Enemies and Control. Elsevier Science B.V., Amsterdam. pdf
  98. Gullan, P.J., Cranston, P.S. and Cook, L.G. (1997) The response of gall-inducing scale insects (Hemiptera: Eriococcidae: Apiomorpha Rübsaamen) to the fire history of mallee eucalypts in Danggali Conservation Park, South Australia. Transactions of the Royal Society of South Australia 121: 137–146. pdf
  99. Gullan, P.J. and Kosztarab, M. (1997) Adaptations in scale insects. Annual Review of Entomology 42: 23–50. pdf
  100. Gullan, P.J. and Strong, K.L. (1997) Scale insects under eucalypt bark: a revision of the Australian genus Phacelococcus Miller (Hemiptera: Coccoidea: Eriococcidae). Australian Journal of Entomology. 36: 229–240. pdf
  101. Melbourne, B.A., Gullan, P.J. and Su, Y.-N. (1997) Interpreting data from pitfall-trap surveys: crickets and slugs in exotic and native grasslands of the Australian Capital Territory. Memoirs of the Museum of Victoria. 56(2): 361–367.
  102. Gullan, P.J. and Stewart, A.C. (1996) A new genus and species of ant-associated coccid (Hemiptera: Coccidae: Myzolecaniinae) from Canthium Lam. (Rubiaceae). Memoirs of the Queensland Museum 39: 307–314.
  103. Hunt, A.J., Gullan, P.J. and Reid, C.A.M. (1996) Chrysomelidae (Coleoptera) and other phytophagous insects in a plantation of black wattle, Acacia mearnsii De Wild., in southeastern Australia. Australian Journal of Entomology 35: 85–92.
  104. Qin, T.K. and Gullan, P.J. (1995) A cladistic analysis of wax scales (Hemiptera: Coccoidea: Coccidae: Ceroplastinae). Systematic Entomology 20: 289–308. pdf
  105. Cranston, P.S. and Gullan, P.J. (1995) Gums and galls. Australian Natural History  24(12): 66–67.
  106. Qin, T.K., Gullan, P.J., et al. (1994) The current distribution and geographical origin of the scale insect pest Ceroplastes sinensis (Hemiptera: Coccidae). Bulletin of Entomological Research 84(4): 541–550. pdf
  107. Gullan, P.J. and Cranston, P.S. (1994) The Insects: An Outline of Entomology. Chapman and Hall, London. 491 pp. (plus 3 reprintings).
  108. Qin, T.K. and Gullan, P.J. (1994) Taxonomy of the wax scales (Hemiptera: Coccidae: Ceroplastinae) in Australia.Invertebrate Taxonomy 8: 923–959.
  109. Campbell, M.H., Holtkamp, R.H., McCormick, L.H., Wykes, P.J., Donaldson, J.F., Gullan, P.J. and Gillespie, P.S. (1994) Biological control of the native shrubs Cassinia spp. using the native scale insects Austrotachardia sp. and Paratachardina sp. (Hemiptera: Kerriidae) in New South Wales.Plant Protection Quarterly 9(2): 64–68. pdf
  110. Gullan, P.J., Buckley, R.C. and Ward, P.S. (1993) Ant-tended scale insects (Hemiptera: Coccidae: Myzolecanium) within lowland rain forest trees in Papua New Guinea. Journal of Tropical Ecology 9: 81–91. pdf
  111. Rozario, S.A., Farrow, R.A. and Gullan, P.J. (1993) The effects of ant attendance on reproduction and survival of Eurymeloides punctata (Signoret) and Eurymela distincta Signoret (Hemiptera: Eurymelidae) on eucalypts. Journal of the Australian entomological Society 32: 177–186.
  112. Qin, T.K. and Gullan, P.J. (1992) A revision of the Australian pulvinariine soft scales (Insecta: Hemiptera: Coccidae).Journal of natural History 26:103–164. pdf1  pdf2
  113. Rozario, S.A., Gullan, P.J. and Farrow, R.A.(1992) Aspects of the biology of Eurymeloides punctata (Signoret) and Eurymela distincta Signoret (Hemiptera: Eurymelidae) feeding on eucalypts.J. Aust. ent. Soc. 31: 317–325.
  114. Gullan, P.J. and Vranjic, J.A. (1991) The taxonomy of the gum tree scales Eriococcus confusus Maskell and E. coriaceus Maskell (Hemiptera: Coccoidea: Eriococcidae).General and Applied Entomology 23: 21–40.pdf
  115. Buckley, R.and Gullan, P. (1991) More aggressive ant species (Hymenoptera: Formicidae) provide better protection for soft scales and mealybugs (Homoptera: Coccidae, Pseudococcidae).Biotropica 23(3): 282–286. pdf
  116. Cranston, P.S., Gullan, P.J. and Taylor, R.W. (1991) Principles and practice of systematics.Ch. 4. pp. 109-124. In: CSIRO.The Insects of Australia. 2nd ed. Melbourne University Press.
  117. Qin, T.K. and Gullan, P.J. (1991) A new species of Kilifia De Lotto from Guizhou, China (Homoptera: Coccidae).Entomotaxonomia 13(1): 21–24.
  118. Bhatti, S. and Gullan, P.J. (1990) New margarodid species (Homoptera: Coccoidea: Margarodidae: Monophlebinae) from New Guinea. Invertebrate Taxonomy 3: 877–911.
  119. Buckley, R.C., Gullan, P.J., Fletcher, M.J. and Taylor, R.W. (1990) New ant-homopteran interactions from tropical Australia.Australian entomological Magazine 17: 57–60.
  120. Qin, T.K. and Gullan, P.J. (1990) The Australian mealybugs (Homoptera: Pseudococcidae) of Xanthorrhoea (Xanthorrhoeaceae). Invertebrate Taxonomy 3: 759–769. pdf
  121. Vranjic, J.A. and Gullan, P.J. (1990) The effect of a sap-sucking herbivore, Eriococcus coriaceus (Homoptera: Eriococcidae), on seedling growth and architecture in Eucalyptus blakelyi. Oikos 59: 157–162. pdf
  122. Gullan, P.J. and Jones, M.G. (1989) A new species of gall-forming coccoid (Insecta: Homoptera: Eriococcidae) from Western Australia. Records of the Western Australian Museum 14: 321–329.
  123. Qin, T.K. and Gullan, P.J. (1989) Cryptostigma Ferris:a coccoid genus with a strikingly disjunct distribution (Homoptera: Coccidae). Systematic Entomology 14: 221–232. pdf
  124. Gullan, P.J., Donaldson, S.R. and Knox, G.A. (1987) Mealybugs (Coccoidea: Pseudococcidae) from the Australian National Botanic Gardens, Canberra. Australian entomological Magazine 14: 49–53. pdf
  125. Gullan, P.J. and Cockburn, A. (1986) Sexual dichronism and intersexual phoresy in gall-forming coccoids.Oecologia (Berlin) 68: 632–634. pdf
  126. Gullan, P.J. (1984a) Froggatt's accession notebooks.Australian entomological Magazine 10: 91–92.
  127. Gullan, P.J. (1984b) A revision of the gall-forming coccoid genus Apiomorpha Rübsaamen (Homoptera: Eriococcidae: Apiomorphinae). Australian Journal of Zoology Supplementary Series, No. 99: 203 pp.
  128. Gullan, P.J. (1984c) A revision of the gall-forming coccoid genus Cylindrococcus Maskell (Homoptera: Eriococcidae).Australian Journal of Zoology 32: 677–690.
  129. Gullan, P.J. (1983) Spiracular structure of adult females of Apiomorpha Rübsaamen (Hemiptera: Coccoidea: Eriococcidae). Journal of the Australian entomological Society 22: 25–29. pdf
  130. Gullan, P.J. (1979) The wax-exuding, cuticular pores of Apiomorpha Rübsaamen (Homoptera: Coccoidea):a light microscopy and scanning electron microscopy study. Australian entomological Magazine 6: 3–9.
  131. Gullan, P.J. (1978) Male insects and galls of the genus Cylindrococcus Maskell (Homoptera: Coccoidea). Journal of the Australian entomological Society 17: 53–61. pdf
  132. Gullan, P.J. (1977) A revision of the genus Anilicus Candèze, with notes on related genera (Coleoptera: Elateridae). Memoirs of the national Museum of Victoria 38: 209–230.

Unrefereed Conference Proceedings, Book Reviews and other publications

  1. Cranston, P.S. & Gullan, P.J. (2014) A dyeing business? Canary cochineal insects. Antenna 38(4): 202–207. pdf
  2. Ouvrard, D., Kondo, T. & Gullan, P.J. (2013) Scale insects: Major Pests and Management. In: Encyclopedia of Pest Management. Taylor and Francis, New York, published online 01 March 2013, pp. 1–4. http://www.tandfonline.com/doi/book/10.1081/E-EPM 
  3. Gullan, P. (2012) Beautiful bird-of-paradise 'flies' and their giant mates. Victorian Entomologist 42(2): 22–25. pdf
  4. Gullan, P. and Kondo, T. (2011) Nomenclator entomologicus. No. 79. (In Portuguese). Informativo da Sociedade Entomológica do Brasil. February 36(1): 3. [A short note on recent changes to the classification relating to Icerya].
  5. Gullan, P.J. and Williams, D.J. (2010) Obituary. Helen May Brookes (3.11.1917-1.1.2008). Entomologica Hellenica 19: 170-172. pdf
  6. Gullan, P.J. and Williams, D.J.(2009) Chris J. Hodgson – much more than an expert coccidologist. pp. 315–317. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal. [THIS IS A REVIEW OF THE CAREER & CONTRIBUTIONS OF Dr C.J. Hodgson]  pdf
  7. Gullan, P.J.(2009) Jan Koteja– an extraordinary evolutionary and revolutionary coccidologist. pp. 305–307. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal. [THIS IS A REVIEW OF THE SCIENTIFIC CONTRIBUTIONS OF THE LATE Dr J. Koteja]  pdf
  8. Gullan, P.J. (2006) Book reviews of A Systematic Catalogue of the Scale Insect Family Margarodidae (Hemiptera: Coccoidea) by Y. Ben-Dov (2005) and A Systematic Catalogue of the Cerococcidae, Halimococcidae, Kermesidae, Micrococcidae, Ortheziidae, Phenacoleachiidae, Phoenicococcidae and Stictococcidae (Hemiptera: Coccoidea) by D.R. Miller, M.E. Gimpel and A. Rung (2005). Systematic Entomology 31: 370.
  9. Cranston, P.S. and Gullan, P.J. (2005) Time flies?  Evolution 59(11): 2492-2494. [Book review of D. Grimaldi's and M.S. Engel's "Evolution of the Insects" ]
  10. Gullan, P.J. (1996) Supervision of research students in biological science. pp. 6-10. In: Supervision of Research Students, Occasional Paper GS96/2, The Graduate School, ANU,Canberra, ACT.
  11. Gullan, P.J. (1996) Book review of Entomology, 2nd ed. by C. Gillott (1995). Aust. J. Ent.35: 196.
  12. Gullan, P.J. (1995) Book review of The Scale Insect Family Coccidae: An Identification Manual to Genera by C.J. Hodgson (1994).Ento. mon. Mag. 131: 186.
  13. Gullan, P.J. (1991) Book review of Margarodidae (Insecta: Hemiptera). Fauna of NewZealand Number 21 by C.F. Morales (1991). N.Z.J. Zool. 18: 451–453.
  14. Gullan, P.J. (1990) Book review of The Scale Insects of the Tropical South Pacific Region. 3 volumes by D.J. Williams& G.W. Watson (1988-1990). Bull. ent. Res. 80: 487–8.
  15. Gullan, P.J. and Buckley, R.C. (1990) Coccids in ants' nests inside live plants in Australia and Papua New Guinea.Proc. ISSIS-VI, Kraków, p. 82.
  16. Gullan, P.J. (1988) The biology of Auloicerya acaciae Morrison and Morrison - an Australian iceryine margarodid.Boll.Lab. Ent. agr. Filippo Silvestri 43 (1986), Suppl.pp. 155–160. pdf
  17. Gullan, P.J.(1987) Book review of Fauna ofNew Zealand, No. 11. Pseudococcidae (Insecta: Hemiptera). J. Aust. ent. Soc. 26: 234.
  18. Gullan, P.J.(1984) Taxonomy and biology of Australian gall-forming Coccoidea.Verh.SIEEC. X., Budapest, pp. 381–383.